Andrena

Pierre Rasmont, Stuart P.M. Roberts, Denis Michez, Oliver Schweiger, Sébastien Patiny, Markus Franzén, Thibaut De Meulemeester, Bodgan Tomozei & Vladimir G. Radchenko
Atlas of the European Bees: genus Andrena

First on line 15.VIII.2013
Updated 6.IX.2013, 16.X.2013, 3.VII.2014

This page is constructed in the framework of the STEP project
Status and Trends of European Pollinators
Coordinator: Simon Geoffrey Potts, University of Reading

STEP Partners to Objective 1 (Document the status and trends of pollinators, map distributions):
University of Mons (Prof. Pierre Rasmont; Denis Michez; Stephanie Iserbyt; Yvan Barbier)
University of Reading (Stuart Roberts)

This first version is mainly based on the Klaus Warncke's manuscript conserved at the Oberösterreichishes Landesmuseum, Linz (Dr. F. Gusenleitner). It also includes a huge number of data sent by several big national and regional databases (see next section for the detail).

Many thanks to Yvan Barbier, Jean-Marc Michalowski, Alain Pauly and Nicolas Vereecken for the pictures.

Pierre Rasmont

Citation:
Rasmont P. Roberts S.P.M., Michez D., Schweiger O., Franzen M., De Meulemeester T., Tomozei B. & Radchenko V. 2013. Atlas of the European Bees: genus Andrena. 1st Edition. STEP Project, Atlas Hymenoptera, Mons, Gembloux. http://www.atlashymenoptera.net/page.aspx?ID=243

Andrena A - C (present page)
Andrena D - K
Andrena L - O
Andrena P - S
Andrena T - Z

Andrena Fabricius, 1775

European bees
Andrena sp. Fabricius, 1775
This map summarizes the data already used. It includes a total of 376,845 data for 598 species.

Main contributions to the maps:

Datasets included 15.IX.2013 (total 376,845 data):
SSIC (Sweden, M. Franzén, B. Cederberg) 115,846 data; BWARS (UK, S.P.M. Roberts) 80,390 data; BDFGM (P. Rasmont & E. Haubruge) 61,987 data; CSCF (Suisse, Y. Gonseth) 48,596 data; UFZ (Germany, O. Schweiger) 41,121 data; Czech Republic (J. Straka) 13,601 data,; FMNH (Finland, J. Paukkunen) 8,449 data; NBDC (Eire, U. FitzPatrick) 3,833 data; NSIC (Norway, F. Odegaard) 1,517 data; Portugal (D. Baldock) 549 data; Waarnemingen.be (Belgium, J. D'Haeselaar) 479 data; others 476 data.

Litterature data: That includes mainly the data from the K. Warncke's manuscript : 46,908 data

We thank the next determinators (first 90%):
Allen G, Allen GW, Amiet F, Archer ME, Artmann-Graf G, Baldock DW, Beavis IC, Bur M, Cheeseborough I, Claessens C, Collins GA, Dicker GHL, Edwards M, Else GR, Falk SJ, Felton JC, Franzen M, Gibbs DJ, Hallin G, Harvey PR, Hellqvist S, Herrmann M, Larsson M, Leclercq J, Lefeber V, Morris RK, Müller A, Neumeyer-Funk R, Norén L, Ødegaard F, Patiny S, Paukkunen J, Petit J, Raekunnas M, Rasmont P, Remacle A, Roberts SPM, Robinson NA, Söderman G, Spooner GM, Steinmann E, Straka J, Strudwick T, Thirion C, Warncke K, Wright IR.

We thank the next recorders (first 90%):
Abenius J, Adamski A, Ahola M, Alexander KNA, Allen G, Allen GW, Amiet F, Andersson H, Andrewes L, Antonsson K, Archer ME, Artmann-Graf G, Bächli G, Baldock DW, Barbier Y, Barone R, Bartsch H, Batelka J , Beavis IC, Bengtsson C, Bengtsson S, Bergsten J, Bergström L, Bernasconi-Ockroy M, Bieri S, Blažej L, Boberg H, Bovey P, Bucher P, Bur M, Burn JT, Butterfield R, Carron G, Cavro, Cederberg B, Chal A, Cheeseborough I, Chittaro Y, Claessens C, Clemons L, Collins GA, Cross IC, Danforth BN, Däniker E, David CT, Davidson AS, Day CD, de Beaumont F, Denton JS, Dicker GHL, Dickson R, Edelsjö J, Edqvist M, Edwards M, Elfving R, Elgar H, Else GR, Ely WA, Erlandsson SO, Falk SJ, Farkac J , Fassotte C, FH Haines, Flint JH, Flodin P, Fordham WJ, Franzen M, Frey-Gessner E, Frisby GE, Gainey P, Gaunitz DVM, Gibbs DJ, Grabs A, Grootaert P , Grosjean J, Guichard KM, Haes ECM, Hallin G, Halstead AJ, Hansson E, Harvey PR, Harwood P, Hawkins RD, Hedin J, Hellqvist S, Herrmann M, Hirt K & H, Hodge PJ, Holmström G, Hrcek J, Ikävalko J, Ingemar Alenäs, Ivarsson T, J Wood, Jacob BJ, Jarman AP, Jensen K, Johansson N, Johansson S, Jones N, Jönsson S, Jukes A, Kaby M, Kakko I, Karhu A, Karlsson Å, Karlsson Bo, Karlsson R, Karlsson T, Kihl A , Kihl S, Knowles A, Koski H , Larsson FK, Larsson K, Lee Rudd, Leinonen R, Longo M, Lönnell N, Lundsten K-E, Macdonald M, Malmqvist A, Mangsbo D, McGee K, Merz B, Mikael Tuneld, Miles SR, Monnerat C, Moretti M, Morgan IK, Morris RK, Müller A, Mura M, Nadig A senior, Neumeyer-Funk R, Nicklas Jansson Län, Nielsen et al, Niemelä P, Nilsson A, Nilsson J, Nilsson LGR, Nilsson SG, Norén L, Ødegaard F, Oertli S, Örjan F, Östrand F, Owens N, Packer L, Patiny S, Paukkunen J, Paul M, Pavett M, Pavett PM, Persson M, Persson S, Persson Vinne T, Peter Nolbrant, Petters MW, Phillips AL, Philp EG, Philpott AJ, Pinchen BJ, Plant cw, Pöyry J, Preisler J, Preisler V, Raekunnas M, Rasmont P, Remacle A, Reser-Rezbanyai L, Roberts SPM, Robinson NA, Roger Pettersson gn, Ronayne C, Saarinen K, Salin T, Saunders E, Schindler M, Schmid W, SJGregory, Sjödin E, Smith MN, Söderman G, Sörensson M, Spooner GM, Steck-Hofmann T, Steinmann E, Stenmark M, Stenmark M M, Straka J, Strudwick T, Svensson S, Tengö J, Terzo M, Tinner-Guler H, Tranefors T, Valkeila E, Verleysen, Webb J Jr, Wedmore RW, Wegelin H, Weibull H, Wolf JP, Wood J, Wright IR, Yeo PF, Zemp F,

P. Rasmont
photo
Andrena abbreviata DOURS,1873
Andrena (Ulandrena) abbreviata DOURS,1873

This taxon includes the next sspp.
ssp. abbreviata s.s. from Crete
ssp. osychniuki Osytshnyuk,1977, from Ukraine, Caucasus, Romania, Bulgaria, former Yugoslavia (Macedonia), Greece, W.Turkey
ssp. abnormis Warncke,1967 from Israel
ssp. dominica Warncke,1975 from S.Turkey
ssp. polemediana Mavromoustakis,1956

Some of these sspp. (osychniuki and polemediana) have been taken as good species by Gusenleitner & Schwarz,2002.
Andrena aberrans EVERSMANN, 1852
Andrena (Taeniandrena) aberrans EVERSMANN, 1852
Andrena abjecta PEREZ,1895
Andrena (Micrandrena) abjecta PEREZ,1895
Andrena abrupta WARNCKE,1967
Andrena (Chlorandrena) abrupta WARNCKE,1967
Andrena acerba WARNCKE,1967
Andrena (Ulandrena) acerba WARNCKE,1967
Andrena aciculata MORAWITZ,1886
Andrena (Aciandrena) aciculata MORAWITZ,1886
Andrena acrana WARNCKE,1967
Andrena (Orandrena) acrana WARNCKE,1967
Andrena adjacens MORAWITZ,1876
Andrena (Poecilandrena) adjacens MORAWITZ,1876
Andrena aegyptiaca FRIESE,1899
Andrena (Chrysandrena) aegyptiaca FRIESE,1899
Andrena aegypticola FRIESE,1922
Andrena (Suandrena) aegypticola FRIESE,1922
Andrena aeneiventris MORAWITZ,1872
Andrena (Aenandrena) aeneiventris MORAWITZ,1872
Andrena aerinifrons DOURS,1873
Andrena (Carandrena) aerinifrons DOURS,1873
Andrena aetherea WARNCKE,1974
Andrena (Suandrena) aetherea WARNCKE,1974
Andrena afrensis WARNCKE,1967
Andrena (Aquandrena) afrensis WARNCKE,1967
Andrena agilissima (SCOPOLI,1770)
Andrena (Aquandrena) agilissima (SCOPOLI,1770)
Andrena agnata WARNCKE,1967
Andrena (Chlorandrena) agnata WARNCKE,1967
Andrena aiderensis OSYTSHNYUK,1993
Andrena (Micrandrena) aiderensis OSYTSHNYUK,1993
Andrena ailisensis OSYTSHNYUK,1993
Andrena (Micrandrena) ailisensis OSYTSHNYUK,1993
Andrena albifacies RADOSZKOWSKI,1874
Andrena (Melandrena) albifacies RADOSZKOWSKI,1874
Andrena albopicta RADOSZKOWSKI,1874
Andrena (Truncandrena) albopicta RADOSZKOWSKI,1874
Andrena albopunctata (ROSSI,1792)
Andrena (Melandrena) albopunctata (ROSSI,1792)
Andrena alchata WARNCKE,1974
Andrena (Truncandrena) alchata WARNCKE,1974
Andrena alfkenella PERKINS,1914
Andrena (Micrandrena) alfkenella PERKINS,1914
Andrena alfkenelloides WARNCKE,1965
Andrena (Micrandrena) alfkenelloides WARNCKE,1965
Andrena alijevi OSYTSHNYUK,1986
Andrena (Euandrena) alijevi OSYTSHNYUK,1986
Andrena alisadra ARIANA,SCHEUCHL,TADAUCHI & GUSENLEITNER,2009
Andrena (Brachyandrena) alisadra ARIANA,SCHEUCHL,TADAUCHI & GUSENLEITNER,2009
Andrena allosa WARNCKE,1975
Andrena (Euandrena) allosa WARNCKE,1975
Andrena alluaudi WARNCKE,1967
Andrena (Chrysandrena) alluaudi WARNCKE,1967
Andrena amicula WARNCKE,1967
Andrena (Aciandrena) amicula WARNCKE,1967
Andrena anatolica ALFKEN,1935
Andrena (Nobandrena) anatolica ALFKEN,1935
Andrena angustior (KIRBY,1802)
Andrena (Ptilandrena) angustior (KIRBY,1802)
Andrena anthrisci BLUETHGEN,1925
Andrena (Micrandrena) anthrisci BLUETHGEN,1925
Andrena antigana PEREZ,1895
Andrena (Simandrena) antigana PEREZ,1895
Andrena apicata SMITH,1847
Andrena (Andrena) apicata SMITH,1847
Andrena apiformis KRIECHBAUMER,1873
Andrena (Plastandrena) apiformis KRIECHBAUMER,1873
Andrena argentata SMITH,1844
Andrena (Leucandrena) argentata SMITH,1844
Andrena argyreofasciata SCHMIEDEKNECHT,1900
Andrena (Thysandrena) argyreofasciata SCHMIEDEKNECHT,1900
Andrena arsinoe SCHMIEDEKNECHT,1900
Andrena (Aciandrena) arsinoe SCHMIEDEKNECHT,1900
Andrena aruana WARNCKE,1967
Andrena (Cryptandrena) aruana WARNCKE,1967
Andrena asiatica FRIESE,1921
Andrena (Nobandrena) asiatica FRIESE,1921
Andrena asperrima PEREZ,1895
Andrena (Aquandrena) asperrima PEREZ,1895
Andrena asperula OSYTCHNJUK,1977
Andrena (Euandrena) asperula OSYTCHNJUK,1977
Andrena assimilis RADOSZKOWSKI,1876
Andrena (Melandrena) assimilis RADOSZKOWSKI,1876
Andrena astica WARNCKE,1967
Andrena (Chlorandrena) astica WARNCKE,1967
Andrena athenensis WARNCKE,1965
Andrena (Nobandrena) athenensis WARNCKE,1965
Andrena atrata FRIESE,1887
Andrena (Parandrenella) atrata FRIESE,1887
Andrena atrotegularis HEDICKE,1923
Andrena (Melandrena) atrotegularis HEDICKE,1923
Andrena avara WARNCKE,1967
Andrena (Avandrena) avara WARNCKE,1967
Andrena azerbaidshanica LEBEDEV,1932
Andrena (Notandrena) azerbaidshanica LEBEDEV,1932
Andrena balcanica WARNCKE,1964
Andrena (Avandrena) balcanica WARNCKE,1964
Andrena barbareae PANZER,1805
Andrena (Melandrena) barbareae PANZER,1805
Andrena barbilabris (KIRBY,1802)
Andrena (Leucandrena) barbilabris (KIRBY,1802)
photo
Andrena bassana WARNCKE,1969
Andrena (Truncandrena) bassana WARNCKE,1969
Andrena bayona WARNCKE,1975
Andrena (Micrandrena) bayona WARNCKE,1975
Andrena bellidis PEREZ,1895
Andrena (Carandrena) bellidis PEREZ,1895
Andrena bengasinensis VON SCHULTHESS,1924
Andrena (Truncandrena) bengasinensis VON SCHULTHESS,1924
Andrena biarmica WARNCKE,1975
Andrena (Micrandrena) biarmica WARNCKE,1975
Andrena bicolor FABRICIUS,1775
Andrena (Euandrena) bicolor FABRICIUS,1775
photo  photo  photo  photo
Andrena bicolorata (ROSSI,1790)
Andrena (Hyperandrena) bicolorata (ROSSI,1790)
Andrena bifida WARNCKE,1967
Andrena (Chlorandrena) bifida WARNCKE,1967
Andrena biguttata FRIESE,1922
Andrena (Ulandrena) biguttata FRIESE,1922
Andrena bilavia OSYTSHNYUK,1994
Andrena (Graecandrena) bilavia OSYTSHNYUK,1994
Andrena bimaculata (KIRBY,1802)
Andrena (Plastandrena) bimaculata (KIRBY,1802)
Andrena binominata SMITH,1853
Andrena (Carandrena) binominata SMITH,1853
Andrena biskrensis PÉREZ, 1895
Andrena (Simandrena) biskrensis PÉREZ, 1895
Andrena bisulcata MORAWITZ,1878
Andrena (Aenandrena) bisulcata MORAWITZ,1878
Andrena blanda PEREZ,1895
Andrena (Poliandrena) blanda PEREZ,1895
Andrena bonasia WARNCKE,1969
Andrena (Aenandrena) bonasia WARNCKE,1969
Andrena boyerella DOURS,1892
Andrena (Chlorandrena) boyerella DOURS,1892

A large part of the data for this species deal with A. leucolippa Pérez, that has been considered as a ssp. of boyerella.
A. boyerella s.s. inhabits N. Africa and likely S. Spain while A. leucolippa lives in Europe. The limits between the distribution of these species is yet unknown.
Andrena braunsiana FRIESE,1887
Andrena (Pallandrena) braunsiana FRIESE,1887
Andrena breviscopa PÉREZ, 1895
Andrena (Simandrena) breviscopa PÉREZ, 1895
Andrena brumanensis Friese,1899
Andrena (Cryptandrena) brumanensis Friese,1899
Andrena bucephala STEPHENS,1846
Andrena (Hoplandrena) bucephala STEPHENS,1846
Andrena bulgariensis WARNCKE,1965
Andrena (Andrena) bulgariensis WARNCKE,1965
Andrena butea WARNCKE,1965
Andrena (Graecandrena) butea WARNCKE,1965
Andrena bytinskii Warncke
Andrena (Poecilandrena) bytinskii Warncke
Andrena caesia WARNCKE,1974
Andrena (Taeniandrena) caesia WARNCKE,1974
Andrena calandra WARNCKE,1975
Andrena (Micrandrena) calandra WARNCKE,1975
Andrena callosa WARNCKE,1967
Andrena (Chlorandrena) callosa WARNCKE,1967
Andrena caneae STRAND,1915
Andrena (Truncandrena) caneae STRAND,1915
Andrena caneibia STRAND,1915
Andrena (Micrandrena) caneibia STRAND,1915
Andrena canohirta (FRIESE,1922)
Andrena (Avandrena) canohirta (FRIESE,1922)
Andrena cantiaca WARNCKE,1975
Andrena (Ulandrena) cantiaca WARNCKE,1975
Andrena capillosa MORAWITZ,1876
Andrena (Euandrena) capillosa MORAWITZ,1876
Andrena capilosella OSYTSHNYUK,1986
Andrena (Euandrena) capilosella OSYTSHNYUK,1986
Andrena caprimulga WARNCKE,1975
Andrena (Lepidandrena) caprimulga WARNCKE,1975
Andrena cara Nurse,1904
Andrena (Carandrena) cara Nurse,1904
Andrena carbonaria sensu lato
Andrena (Plastandrena) carbonaria sensu lato
=Andrena pilipes Fabricius, 1781 + Andrena nigrospina Thomson,1872
photo
Andrena caroli PEREZ,1895
Andrena (Campylogaster) caroli PEREZ,1895
Andrena caspica MORAWITZ,1886
Andrena (Poliandrena) caspica MORAWITZ,1886

The true caspica MORAWITZ is a species from the Caucasian region.
The map here represents the distribution of the near taxa A. unicincta and A. westensis.
A. westensis Warncke 1965 lives in Greece and as been described as an unicincta ssp.
A. unicincta Friese 1899 has been described from Israel.

This map should be deeply revised.

Andrena caudata WARNCKE,1966
Andrena (Avandrena) caudata WARNCKE,1966
Andrena cervina WARNCKE,1975
Andrena (Micrandrena) cervina WARNCKE,1975
Andrena chaetogastra PITTIONI,1950
Andrena (incertae sedis) chaetogastra PITTIONI,1950
Andrena chalcogastra BRULLE,1839
Andrena (Taeniandrena) chalcogastra BRULLE,1839
Andrena chelma WARNCKE,1975
Andrena (Aciandrena) chelma WARNCKE,1975
Andrena chersona WARNCKE,1972
Andrena (incertae sedis) chersona WARNCKE,1972
Andrena chrysopus PEREZ,1903
Andrena (Euandrena) chrysopus PEREZ,1903
Andrena chrysopyga SCHENCK,1853
Andrena (Zonandrena) chrysopyga SCHENCK,1853
Andrena chrysosceles (KIRBY,1802)
Andrena (Notandrena) chrysosceles (KIRBY,1802)
photo  photo  photo  photo
Andrena ciconia WARNCKE,1975
Andrena (Poecilandrena) ciconia WARNCKE,1975
Andrena cineraria (L.,1758)
Andrena (Melandrena) cineraria (L.,1758)
photo  photo  photo  photo
Andrena cinerea BRULLE,1832
Andrena (Chlorandrena) cinerea BRULLE,1832
Andrena cinereophila WARNCKE,1965
Andrena (Chlorandrena) cinereophila WARNCKE,1965
Andrena cinnamonea WARNCKE,1975
Andrena (Simandrena) cinnamonea WARNCKE,1975
Andrena clarkella (KIRBY,1802)
Andrena (Andrena) clarkella (KIRBY,1802)
photo  photo  photo
Andrena clusia WARNCKE,1966
Andrena (Hoplandrena) clusia WARNCKE,1966
Andrena clypella STRAND,1921
Andrena (Chlorandrena) clypella STRAND,1921
Andrena coitana (KIRBY,1802)
Andrena (Stenomelissa) coitana (KIRBY,1802)
Andrena colletiformis MORAWITZ,1874
Andrena (Brachyandrena) colletiformis MORAWITZ,1874
Andrena colonialis MORAWITZ,1886
Andrena (Euandrena) colonialis MORAWITZ,1886
Andrena combaella WARNCKE,1966
Andrena (Ulandrena) combaella WARNCKE,1966
Andrena combinata (CHRIST,1791)
Andrena (Simandrena) combinata (CHRIST,1791)
Andrena compta LEPELETIER,1841
Andrena (Nobandrena) compta LEPELETIER,1841
Andrena comptaeformis GUSENLEITNER & SCHWARZ,2000
Andrena (Nobandrena) comptaeformis GUSENLEITNER & SCHWARZ,2000
Andrena comta EVERSMANN,1852
Andrena (Melandrena) comta EVERSMANN,1852
Andrena concinna SMITH,1853
Andrena (Ulandrena) concinna SMITH,1853
Andrena congruens SCHMIEDEKNECHT,1883
Andrena (Simandrena) congruens SCHMIEDEKNECHT,1883
Andrena corax WARNCKE,1967
Andrena (Poliandrena) corax WARNCKE,1967
Andrena cordialis MORAWITZ,1878
Andrena (Cordandrena) cordialis MORAWITZ,1878
Andrena coromanda WARNCKE,1975
Andrena (Melittoides) coromanda WARNCKE,1975
Andrena corssubalpina THEUNERT,2007
Andrena (Micrandrena) corssubalpina Theunert,2007

This recently described species is endemic to Corsica. It lives on rocky slopes at the highest level of subalpine to alpine level. It forages on the endemics plants Ligusticum corsicum (Umbelliferae) and Cerastium soleirolii (Caryophyllaceae) (Theunert, 2007, 2012).
Andrena crassana WARNCKE,1965
Andrena (Poecilandrena) crassana WARNCKE,1965
Andrena creberrima PEREZ,1895
Andrena (Zonandrena) creberrima PEREZ,1895
Andrena crispa WARNCKE,1975
Andrena (Parandrenella) crispa WARNCKE,1975
Andrena cubiceps FRIESE,1914
Andrena (Cubiandrena) cubiceps FRIESE,1914
Andrena cubicepsella WARNCKE,1975
Andrena (Cubiandrena) cubicepsella WARNCKE,1975
Andrena cuneata WARNCKE,1974
Andrena (Carandrena) cuneata WARNCKE,1974
Andrena curiosa (MORAWITZ,1878)
Andrena (Melittoides) curiosa (MORAWITZ,1878)
Andrena curtula PEREZ,1903
Andrena (Micrandrena) curtula PEREZ,1903
Andrena curvana WARNCKE,1965
Andrena (Notandrena) curvana WARNCKE,1965
Andrena curvungula THOMSON,1870
Andrena (Lepidandrena) curvungula THOMSON,1870
Andrena cussariensis MORAWITZ,1866
Andrena (Melandrena) cussariensis MORAWITZ,1866
Andrena cyanomicans PEREZ,1895
Andrena (Suandrena) cyanomicans PEREZ,1895
Andrena cypria PITTIONI,1950
Andrena (Cordandrena) cypria PITTIONI,1950
Andrena cypricola MAVROMOUSTAKIS,1952
Andrena (Plastandrena) cypricola MAVROMOUSTAKIS,1952

References

Ariana A., Scheuchl E., Tadauchi O. & Gusenleitner F. 2009. A taxonomic revision of the subgenus Andrena (Brachyandrena) (Hymenoptera: Andrenidae). Zootaxa. 2281: 21–39.
see also:http://www.mapress.com/zootaxa/2009/f/zt02281p039.pdf
Blank S.M. & Kraus M. 1994. The nominal taxa described by K. Warncke and their types (Insecta, Hymenoptera, Apoidea). Linzer biol. Beitr. 26(2): 665-761.
Dylewska M. 1987. Die Gattung "Andrena" Fabricius ("Andrenidae", "Apoidea") in Nord-und Mitteleuropa. Acta Zoologica Cracoviensia, 30(12): 359-708.
Gogala A. 2011. Some interesting notes on the Andrena species in Slovenia (Hymenoptera: Andrenidae). Acta Entomologica Slovenica, 19(1): 29–35.
see also:http://www2.pms-lj.si/pdf/Acta/AES_19-1_3_GOGALA.pdf
Grünwaldt W., Osytshnyuk A.Z. & Scheuchl E. 2005. Neue Andrena-Arten aus der Paläarktis (Hymenoptera: Apidae: Andreninae). Entomofauna, 26(19): 349-368.
Gusenleitner F. & Schwarz M. 2002. Weltweite Checkliste der Bienengattung Andrena mit Bemerkungen und Ergänzungen zu paläarktischen Arten (Hymenoptera, Apidae, Andreninae, Andrena). Entomofauna suppl. 12: 1-1280.
see also:http://www.zobodat.at/pdf/ENTS_S12_0001-1280.pdf
Kratochwil A. & Scheuchl E. 2013. Andrena (Micrandrena) dourada nov.sp. from Porto Santo, Madeira Archipelago, Portugal. Linzer biol. Beitr. 45(1): 755-774.
see also:http://www-old.biologie.uni-osnabrueck.de/Oekologie/pdf%20kratochwil/2013_112.pdf
Osytchnjuk A.Z. 1978. Opredelitel' nasekomykh evropejskoj chast SSSR. Tom III. Pereponchatokrylye pervaya chast. Apoidea Andrenidae (ed. K.B. Gorodkov). Akademia Nauk SSSR, Leningrad, pp. 315- [in Russian].
Osytshnjuk A.Z. 1977. Fauna Ukrayini. Tom 12. Bdzholini. Viduck 5. Bdzholi-Andrenidi. Akademiya Nauk Ukrains'koj RSR, Institut Zoologii, Kiev, 328 pp.
Osytshnjuk A.Z. (traduction Kut R.). 1993. Clés de détermination des insectes de la partie européenne de l'U.R.S.S. Tome III. Hyménoptères. Première partie. Extrait: Superfamille Apoidea (apoïdes). Andrenidae. Mémoire de traduction , Université de Mons-Hainaut,EII, 199 pp. [en français].
Osytshnjuk A.Z., Romasenko L., Banaszak J. & Cierzniak T. 2005. Andreninae of the Central and Eastern Palaearctic. Part 1. Polish Entomological Society, Poznan, Bydgoszcz, 235 p.
Osytshnjuk A.Z., Romasenko L., Banaszak J. & Motyka E. 2008. Andreninae of the Central and Eastern Palaearctic. Part 2. Polish Entomological Society, Poznan, Bydgoszcz, 234 p.
Patiny S. & Terzo M. 2010. Catalogue et clé des sous-genres et espèces du genre Andrena de Belgique et du nord de la France (Hymenoptera, Apoidea). Laboratoire de Zoologie, Université de Mons, 39 p.
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